Richard Axel , 1946-, American pathologist and biochemist, b. New York, N.Y., M.D. Johns Hopkins, 1970. A professor at Columbia from 1978, Axel was awarded, with Linda B. Buck , the 2004 Nobel Prize in Medicine or Physiology for elucidating the human olfactory system and demonstrating how olfactory receptors are encoded in the nose. After working together on the discovery of odorant receptors, the two worked in parallel in different labs in the 1990s and went on to learn how the brain organizes signals from those receptors to ...

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