Wood charcoal from Santorini (Thera): new evidence for climate, vegetation and timber imports in the Aegean Bronze Age.(Research)(Greek Islands archaeobotany studies)

Bronze Age climate in the Aegean

Palaeoclimatic reconstructions for the Bronze Age Aegean based on pollen analytical investigations from several sites in mainland Greece (Bottema 1974, 1982, 1990) have suggested climate conditions drier than at present, with a reversal to a wetter climate favouring woodland expansion and arboriculture (including the intensive cultivation of olive trees) occurring only towards the end of the Bronze Age and the beginning of the historical periods (c. 3200/2700 uncal. BP, c.1500/1100 cal. BC). More recent pollen analyses from Crete have refined this general model, indicating contrasting patterns of vegetation change with the Greek mainland, ...

<5mm was in the range of 1-5 for each 
sample). 
 
Taxa Wood anatomical descriptions 
 
Pinus (halepensis/   TS: early -- latewood transition abrupt, latewood 
brutia type) part relatively short. Resin canals present; RLS: 
 Ray tracheids present, occasionally with 
 conspicuously dentate walls. Ray parenchyma cells 
 with pinoid pits. Crossfields with 1-2(3) pits; 
 TLS: Resin canals present. Rays 7-8 cells high, 
 occasionally higher (up to 15 cells) 
 
Cedrus libaniTS: early -- latewood transition mostly gradual, 
 latewood part relatively short. Traumatic resin 
 canals occasionally present in tangential rows at 
 the ring boundary; RLS: Tracheids bearing mostly 
 uniseriate pits with scalloped tori (Fig. 3), the 
 latter particularly visible in the earlywood. Ray 
 tracheids present, uniseriate, thin-walled, with 
 irregular borders. Ray parenchyma cells thick- 
 walled, with taxodioid pits in the earlywood and 
 mostly piceoid pits in the latewood. Crossfields 
 with 1-4 pits; TLS: Resin canals present (TLS 
 surface too narrow for a reliable estimation of 
 ray height) 
 
Cupressaceae TS: early -- latewood transition gradual, latewood 
 part short. Resin canals absent; RLS: Tracheid 
 pits uniseriate. Ray tracheids absent. Transverse 
 ray walls rather thick, smooth. Tangential walls 
 smooth, occasionally nodular. Ray parenchyma 
 with cypressoid pits. Crossfields with 1-2(3) 
 pits; TLS: (TLS surface too narrow for a reliable 
 estimation of ray height) 
 
Taxus baccataTS: early-latewood transition gradual, earlywood 
 cells thick-walled. Resin canals absent; RLS: 
 Tracheid pits uniseriate. Conspicuous helical 
 thickenings on tracheid walls. Rays pits 
 cypressoid. Ray tracheids absent; TLS: (TLS 
 surface too narrow for a reliable estimation of 
 ray height) 
 
Olea TS: Wood diffuse porous, growth rings mostly 
 indistinct. Pores thick-walled, arranged in short 
 radial multiples of 2-4(5), sometimes in 
 clusters, occasionally solitary, dense. 
 Parenchyma frequent, paratracheal-vasicentric, 
 occasionally confluent; RLS: Perforation plates 
 simple. Rays heterogeneous, with a central 
 proportion of strongly procumbent cells and 
 1-2(3) rows of square and/or upright marginal 
 cells. Inter-vessel and ray-vessel pits numerous, 
 alternate, rounded, small. Librifonn fibres 
 present. Gummy deposits occasionally present; 
 TLS: Rays uni- to bi(3)-seriate up to 10(12) 
 cells high (uniseriate rays rare, short) 
 
Punica granatum  TS: Wood diffuse porous, growth rings mostly 
 indistinct, when present undulating. Pores 
 solitary, in short radial multiples of 2(3-4) or 
 clusters. Parenchyma infrequent; RLS: Perforation 
 plates simple. Rays conspicuously heterogeneous, 
 with a short central portion of procumbent cells 
 and numerous rows of square and upright marginal 
 cells. Inter-vessel and vessel-ray pits vestured, 
 alternate. Libriform fibres present, septate. 
 Gummy deposits present; TLS: Rays uni- to 
 biseriate 
 
Arbutus  TS: Wood diffuse porous, growth rings distinct. 
 Pores mostly angular, solitary, in short radial 
 multiples (2-3) or clustered, numerous. 
 Parenchyma mostly apotracheal, occasionally 
 paratracheal (infrequent); RLS: Perforation 
 plates simple. Rays heterogeneous, with a central 
 portion of strongly procumbent cells and one row 
 of upright marginal cells. Inter-vessel pits and 
 vessel-ray pits with slit-like apertures. Fibre 
 tracheids with bordered pits and (infrequent) 
 septate fibres present. Conspicuous helical 
 thickenings present on vessel members and fibre 
 tracheids; TLS: Rays bi- to 3(4)-seriate 
 
MaloideaeTS: Wood diffuse porous. Growth rings distinct. 
 Pores solitary; RLS: Perforation plates simple. 
 Rays homogeneous to slightly heterogeneous with 
 one row of square marginal cells. Fibre-tracheids 
 present. Helical thickenings present on vessel 
 members and tracheids. Vessel-ray pits simple 
 rounded, enlarged; TLS: Rays mostly biseriate 
 
Quercus (type 1 -TS: Wood ring porous, growth rings narrow, 
deciduous)   distinct. Early wood pores solitary, arranged in 
 a single row, large. Latewood pores solitary, in 
 dendritic to radially oblique arrangement. Broad 
 multiseriate rays conspicuous; RLS: Perforation 
 plates simple. Rays homogeneous. Libriform fibres 
 and vasicentric tracheids present. Vasicentric 
 tracheids abundant, forming the greater part of 
 ground tissue. Apertures of vessel-ray pits 
 simple, enlarged, round to horizontally elongate; 
 TLS: Rays uni- and multiseriate, the latter very 
 broad (>

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