Article: Exotic earthworm invasion increases soil carbon and nitrogen in an old-growth forest in southern Quebec.

Abstract: To test whether invasion of exotic earthworms affects soil carbon (C) and nitrogen (N), we sampled the litter and upper mineral soil (to 30 cm) at a series of sites varying in their earthworm populations in an old-growth beech-maple forest at Mont St. Hilaire, southern Quebec. We measured earthworm abundance and biomass using hand-sorting and chemical extraction (allyl isothiocyanate) methods. They gave similar results, though there was evidence of size and species-specific biases. Abundance and biomass of the earthworms ranged from <10 to>100 earthworms x [m.sup.-2] and from <10 to 125 g x [m.sup.-2], respectively, and were correlated with distance from a nearby lake (negatively) and soil pH (positively). The presence of earthworms was associated with a decrease in the mass and thickness and an increase in the C/N quotient of the litter layer. There were no significant changes in C and N mass of the mineral soil between 0 and 10 cm, but the underlying layers (10-20 and 20-30 cm) in sites with>10 earthworms x [m.sup.-2] showed significantly (p <10 to>10 earthworms x [m.sup.-2] may increase ...

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