Article: Our thumbs.

The hand without a thumb is at worst nothing but an animated spatula * and at best a pair of forceps whose points don't meet properly.--John Napier (1)

ORIGINS

Our earliest known ancestor is the fish Rhipidistia (2), which has been extinct for 230 million years. Its pectoral fin is the precursor of our hand, which has 4 fingers, each with 3 phalanges, and a thumb with only 2 phalanges.

It is known that almost without exception, the first ray of mammals, reptiles and amphibians, and humans had 2 phalanges. Galen (AD 165) thought the bones of the thumb were really 3 phalanges and that it lacked a metacarpal. Vesalius in 1543 agreed, saying that ...

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