Article: Mario Capecchi: The man who changed our world

The genetics research that won Mario Capecchi a share of this year's Nobel Prize for medicine may well help to define the science of the 21st century.

But the man himself was marked, in extraordinary ways, by the turbulent history of the century before.

Mr Capecchi's grandfather, a German archaeologist, was accidentally gunned down by his own men during the First World War. His father, an Italian aviator, perished in the Second World War. He himself spent that war destitute in northern Italy after his American mother was arrested and sent to the Dachau concentration camp - a survival tale all the more remarkable for the fact that he was just four years old when his mother was taken away.

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